Query: NC_009776:707785 Ignicoccus hospitalis KIN4/I, complete genome
Lineage: Ignicoccus hospitalis; Ignicoccus; Desulfurococcaceae; Desulfurococcales; Crenarchaeota; Archaea
General Information: Ignicoccus hospitalis Kin4/I was isolated from gravel obtained from the shallow marine hydrothermal system of the Kolbeinsey Ridge, north of Iceland. Hyperthermophilic archaeon. Ignicoccus hospitalis, an anaerobic, hyperthermophilic, chemoautolithotroph, uses hydrogen to reduce elemental sulfur and produce hydrogen sulfide. This organism is also moderately halophilic and acidophilic. Ignicoccus hospitalis is the host organism for Nanoarchaeum equitans.
Subject: NC_011961:528837 Thermomicrobium roseum DSM 5159 plasmid unnamed, complete sequence
Lineage: Thermomicrobium roseum; Thermomicrobium; Thermomicrobiaceae; Thermomicrobiales; Chloroflexi; Bacteria
General Information: Thermomicrobium roseum DSM 5159 was isolated from Yellowstone National Park, USA. Obligate thermophile with unusual cell wall structure. Thermomicrobium roseum is a red-pigmented, rod-shaped, Gram-negative extreme thermophile that possesses both an atypical cell wall composition and an unusual cell membrane that is composed entirely of long-chain 1,2-diols. Analyses of environmental sequences from hot spring environments show that T.roseum displays a low quantity but ubiquitous presence in top layers of microbial mats. Few standard housekeeping genes are found on the megaplasmid, however, it does encode a complete system for chemotaxis including both chemosensory components and an entire flagellar apparatus. T. roseum oxidizes CO aerobically, making it the first thermophile known to do so. In addition, is is propose that glycosylation of its carotenoids plays a crucial role in the adaptation of the cell membrane to this bacterium's thermophilic lifestyle. Because T. roseum is a deep-branching member of this phylum, eventhough this species is not photosynthetic, analysis of the genome provides some insight into the origins of photosynthesis in the Chloroflexi.