Query: NC_016894:2809125 Acetobacterium woodii DSM 1030 chromosome, complete genome
Lineage: Acetobacterium woodii; Acetobacterium; Eubacteriaceae; Clostridiales; Firmicutes; Bacteria
General Information: Acetobacterium woodii is a Gram positive, motile, strict anaerobic, acetogenic bacterium, that relies on Na+ as coupling ion in bioenergetic reactions. The organism can use a wide range of substrates, such as sugars, alcohols, methoxylated aromatic acids or C1 compounds. Electrons derived from these electron donors are used in the Wood-Ljungdahl-pathway where the organism fixes CO2 and produces acetate. The pathway of CO2-fixation is coupled to energy conservation via a chemiosmotic mechanism, one enzyme that seems to be involved is the Rnf complex. The produced Na+ gradient can be used to drive ATP-synthesis or flagella rotation. The ATP synthase is a member of the F1FO class of enzymes and has an unusual hybrid rotor. Can use alternative electron acceptors like the lignin degradation product caffeate.
Subject: NC_002932:296557 Chlorobium tepidum TLS, complete genome
Lineage: Chlorobaculum tepidum; Chlorobaculum; Chlorobiaceae; Chlorobiales; Chlorobi; Bacteria
General Information: This green-sulfur bacterium is a thermophile and was isolated from a New Zealand high-sulfide hot spring. Photosynthetic thermophile. Chlorobium tepidum is a member of the green-sulfur bacteria. It has been suggested that the green-sulfur bacteria were among the first photosynthetic organisms since they are anaerobically photosynthetic and may have arisen early in the Earth's history when there was a limited amount of oxygen present. This organism utilizes a novel photosynthetic system, and harvests light energy using an unusual organelle, the chlorosome, which contains an aggregate of light-harvesting centers surrounded by a protein-stabilized galactolipid monolayer that lies at the inner surface of the cytoplasmic membrane. Unlike many other photosynthetic organisms, the green-sulfur bacteria do not produce oxygen and tolerate only low levels of the molecule. This organism also fixes carbon dioxide via a reverse tricarboxylic acid cycle, using electrons derived from hydrogen or reduced sulfur to drive the reaction, instead of via the Calvin cycle like many other photosynthetic organisms.