Query: NC_017249:8483159 Bradyrhizobium japonicum USDA 6, complete genome Lineage: Bradyrhizobium japonicum; Bradyrhizobium; Bradyrhizobiaceae; Rhizobiales; Proteobacteria; Bacteria General Information: This nitrogen-fixing bacterium develops a symbiotic relationship with the soybean plant Glycine max and is related to other N2-fixing Rhizobia which are symbiotic with legumes. The bacterium establishes itself in a root nodule which provides a protective environment for the organism to live while the bacterium provides the plant cell with nitrogen. This is an agriculturally important symbiotic relationship as it obviates the need for expensive and environmentally damaging fertilizer.Genes that code for proteins involved in root nodulation are carried on the chromosome. The production of the nodulation signal, lipochitin, is directed by genes which are turned on in the presence of plant flavonoid compounds. The bacteria are endocytosed into a cortical cell, and are enclosed within a membrane bound organelle termed the symbiosome.
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General Information: Soil bacterium producing an antituberculosis agent. The characteristic earthy smell of freshly plowed soil is actually attributed to the aromatic terpenoid geosmin produced by species of Streptomyces. There are currently 364 known species of this genus, many of which are the most important industrial producers of antibiotics and other secondary metabolites of antibacterial, antifungal, antiviral, and antitumor nature, as well as immunosuppressants, antihypercholesterolemics, etc. Streptomycetes are crucial in the soil environment because their diverse metabolism allows them to degrade the insoluble remains of other organisms, including recalcitrant compounds such as lignocelluloses and chitin. Streptomycetes produce both substrate and aerial mycelium. The latter shows characteristic modes of branching, and in the course of the streptomycete complex life cycle, these hyphae are partly transformed into chains of spores, which are often called conidia or arthrospores. An important feature in Streptomyces is the presence of type-I peptidoglycan in the cell walls that contains characteristic interpeptide glycine bridges. Another remarkable trait of streptomycetes is that they contain very large (~8 million base pairs which is about twice the size of most bacterial genomes) linear chromosomes with distinct telomeres. These rearrangements consist of the deletion of several hundred kilobases, often associated with the amplification of an adjacent sequence, and lead to metabolic diversity within the Streptomyces group. Sequencing of several strains of Streptomyces is aimed partly on understanding the mechanisms involved in these diversification processes.